Ben Stiller

Alex (voice)

Chris Rock

Marty (voice)

David Schwimmer

Melman (voice)

Jada Pinkett Smith

Gloria (voice)

Sacha Baron Cohen

King Julien (voice)

Cedric the Entertainer

Maurice (voice)

Andy Richter

Mort (voice)

Tom McGrath

Skipper / Fossa / Panicky Man on Subway (voice)

Christopher Knights

Private (voice)

Chris Miller

Kowalski (voice)

Conrad Vernon

Mason (voice)

Eric Darnell

Zoo Announcer / Lemur #1 / Fossa / Subway Car Announcer (voice)